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    <p>Reason for this behaviour is <a href="http://msdn.microsoft.com/en-us/library/system.windows.controls.contentpresenter.recognizesaccesskey%28v=vs.100%29.aspx" rel="noreferrer">ContentPresenter.RecognizesAccessKey</a> which is by default <code>true</code> for <code>datagrid header contentPresenter</code>. If this property is true, it uses <code>AccessText</code> in text of header.</p> <p>Workaround for this is to hook to <code>AutoGeneratingColumn</code> event of dataGrid and replace all single underscore with double underscores. Sample is shown below - </p> <pre><code>&lt;DataGrid AutoGeneratingColumn="DataGrid_AutoGeneratingColumn"/&gt; </code></pre> <p>and in event handler -</p> <pre><code>private void DataGrid_AutoGeneratingColumn(object sender, DataGridAutoGeneratingColumnEventArgs e) { e.Column.Header = e.Column.Header.ToString().Replace("_", "__"); } </code></pre> <hr> <p>Also you can achieve that by <code>overriding ContentTemplate</code> of <code>DataGridColumnHeader</code>. Use <code>TextBlock</code> inside it which does not recognizes access key from a text.</p> <pre><code>&lt;DataGrid ItemsSource="{Binding Objects}"&gt; &lt;DataGrid.ColumnHeaderStyle&gt; &lt;Style TargetType="{x:Type DataGridColumnHeader}" BasedOn="{StaticResource {x:Type DataGridColumnHeader}}"&gt; &lt;Setter Property="ContentTemplate"&gt; &lt;Setter.Value&gt; &lt;DataTemplate&gt; &lt;TextBlock Text="{TemplateBinding Content}" HorizontalAlignment="Center"/&gt; &lt;/DataTemplate&gt; &lt;/Setter.Value&gt; &lt;/Setter&gt; &lt;/Style&gt; &lt;/DataGrid.ColumnHeaderStyle&gt; &lt;/DataGrid&gt; </code></pre>
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